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3. Begonia heliantha Tebbitt Figs 16, 17A Edinburgh Journal of Botany 73 (1): 145 (Tebbitt 2016). – Type : PERU – Puno Region: Prov. Sandia • Entre Sandia y Tambopata; 1700–2200 m a.s.l.; [14°15′ S, 69°25′ W]; 7 Aug. 1965; J.C. Vargas Calderón 16417ª ; holotype: US [US01269467]; isotype: CUZ [2]. Tebbitt (2020: 69). Etymology The epithet derives from the Greek words ‘ helios ’ and ‘ anthos ’, meaning ‘sun’ and ‘flower’. This references the bright yellow, ‘sun-like’ flowers of the species. Specimens examined PERU – Puno Region: Prov. Sandia • ca 17 km on road below Sandia; 14°13.684′ S, 69°24.736′ W; ca 1600 m a.s.l.; 16 Feb. 2002; R.T. Pennington, T.D. Pennington & A. Daza 1113 ; E [E00274868], K, MOL. Description Caulescent, tuberous herb, to 20 cm high. Tuber ellipsoid, 1.1–2.5 × 1.1–2.5 cm, with one growing point. Stem erect, unbranched; internodes to 3.1 cm long, to 2 mm thick, succulent, red, sparsely pubescent. Stipules persistent, ovate, 1–3.5 × 0.5–2 mm, apex acute, aristate, opaque, brown, glabrous, margin fimbriate, ciliate. Leaves 3–6, alternate, basifixed; petiole 1.7–5 cm long, red, densely glandular-pubescent; blade asymmetric, ovate, to 10 × 5 cm, succulent, apex acute to acuminate, base obliquely cordate, basal lobes not overlapping, sinus to 15 mm deep, margin lacking lobes or shortly-lobed, lobes triangular-crenate, dentate, ciliate, upper surface green with paler veins, sometimes flushed black between the veins, moderately pubescent, lower surface pale green, densely pubescent on the major veins, moderately pubescent on the lamina, veins palmate but with 1 primary vein, 7–9 veined from the base, 1–3 secondary veins on the larger side, 1–2 on the smaller side. Inflorescences 1–3, bisexual, axillary, erect, cymose, with up to 2 branches, bearing up to 3 staminate flowers and 2 pistillate flowers, protandrous; peduncle to 16 cm long, red, moderately glandular-pubescent, bracts persistent, obovate to ovate, 1.25–6 × 0.5–4 mm, translucent, brown, glabrous, apex acute to rounded, margin ... |